A pet dog chameleon (Johnny Depp) becomes stranded in the Mojave Desert after his terrarium drops from his owners' automobile by crash. He meets an armadillo called Roadkill (Alfred Molina) who is seeking the mystical "Spirit of the West" and routes the dry chameleon to discover water at a community called Dirt. While roaming the desert, he narrowly avoids being eaten by a vicious red-tailed hawk and has a surreal headache before fulfilling the desert iguana Beans (Isla Fisher), a herdsman's daughter, who takes the chameleon to Dirt, an Old West community populated by desert animals.
Making use of blowing and improvisation to fit in, the chameleon offers himself to the townsfolk as a difficult drifter named Rango. He quickly contravenes of outlaw Gila beast Bad Bill (Ray Winstone), but avoids a shootout when Costs is scared off by the hawk's return. Rango is chased by the hawk until he unintentionally knocks down a vacant water tower which squashes the killer. In reaction, the community mayor, a senior tortoise, (Ned Beatty) assigns Rango the brand-new constable. Meanwhile, the townsfolk fret that when it comes to the hawk dead, the gunslinger Rattlesnake Jake (Bill Nighy), that was afraid of the hawk, will certainly return.
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